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Hivory Towers is the main HUB world found in Yooka-Laylee. It is a factory and business owned by the game's main antagonist, Capital B. There are many rooms within the factory, some which lead to the Grand Tomes. The tomes are magical books that each contain a different world.
The Hivory Towers total collectibles screen includes collectibles from Shipwreck Creek as well.
Hivory Towers is a play-on-words with the phrase "Ivory Towers" and the word "hive." "Ivory Towers" refers to a state of seclusion to practice knowledge, with the large number of books referring to knowledge. Hive alludes to the fact that bees live in hives, since Capital B is a bee.
Note: Everytime the player collects one Pagie in a new world, Trowzer will relocate to the next location in Hivory Towers. For example, if the player retrieves a Pagie in Tribalstack Tropics, he will move to the second hub area near Glitterglaze Glacier.
Trowzer is to the left side coming into Hivory Towers near the entrance. Trowzer will teach the Reptile Roll here.
These Pagies do not appear until the player has earned at least 1 Pagie from Tribalstack Tropics.
The Wrecked Crow's Nest: A wrecked crow's nest located on the top of a hill contains a Pagie flying in a circle. The player can access this area by using the Reptile Roll on a broken red and green ramp that leads to the hills that eventually lead to the wrecked crow's nest
Helping Blasto: On the left side (facing away from Hivory Towers) of Shipwreck Creek, an uphill path can be found that leads to Blasto. By using the Buddy Slam ability on Blasto, he will cough out a Pagie for the player to collect.
Top of the Golden Capital B Statue: The first Pagie in order to progress through the game is found on top of the Golden B statue in the entrance room of Hivory Towers. The player must jump across a number of obstacles and platforms in order to reach the top.
Extinguish the Fire: At the base of the pathway to the second room, there is a fire with a Pagie just behind it. Use the Splashberry bush located in the crates surrounded by water just behind the statue.
Entrance to the Tribalstack Tropics Grand Tome: In between the main entrance of Hivory Towers and the Tribalstack Tropics Grand Tome is a room with a Pagie in a Cagie. To open the Cagie, set fire to the torches in the order P-A-G-I-E using Slurp Shot.
Fiery Pipe: Above the Tribalstack Tropics Grand Tome, there is a pipe that the duo can climb next to. Use Reptile Rush to break the glass. Laylee must use her Sonar Shield to protect the duo from the flames.
Pagie inside the Arcade Machine: Inside of Rextro's arcade room there is a giant arcade machine in the back with a Pagie inside. Break the glass using Sonar 'Splosion and collect the Pagie.
Pagie in the Pipes: Just before the Glitterglaze Glacier Grand Tome, there is a machine with a set of clear pipes, inside the one that has a gap, there is a Pagie cycling through it. Simply climb your way up to collect it.
Pipe Climb: Directly above the Glitterglaze Glacier Grand Tome there are pipes covered in snow that can be climbed to reach a Pagie.
Slippery Slope: In the room before the Glitterglaze Glacier Grand Tome, use Slurp State on the Bee Hive next to the Corplet Security, and continue past the Grand Tome there will be a slope that leads to a Pagie.
Glass Maze: To the right of the room, there is a bookshelf covered in glass. at the first section, go through the left door. Climb onto the books to reach the section above then enter the right door. Finally, walk left until you walk into another section of the bookcase and go through the right door to the Pagie.
Across the wind turbines: To get past the wind tunnel, continuously use Reptile Rush.
Balloon Challenge: In the tunnel where Yooka and Laylee must fly through, near the start of the tunnel is another tunnel near the very top that has butterflies around it that the duo can enter into. In it is a Pagie in a Cagie. Shoot the balloons using Laylee's Sonar Shot ability to reveal the letters P-A-G-I-E and collect these letters to open the Pagie in a Cagie.
Slide Challenge: Slide down into the Hoops within 39 seconds.
The following Quills are located in Shipwreck Creek.
Hivory Towers Entrance Gate (1): A chest with a single quill is located next to the Hivory Towers Entrance gate
Big Tree (1): Next to the Batship Crazy ship is a big brown lone tree by an edge. Next to it is a treasure chest with a one quill.
Entrance to the Ship (1): On the second floor of the entrance to the ship is a treasure chest past two breakable barrels. The chest contains one quill.
Ship's Quarterdeck (1): On the Quarterdeck (the top floor with the helm) is a treasure chest that contains a quill inside of it.
Red & Green Ramp to the Hills (1): A slightly-broken red and greed ramp lays on a wall with a treasure chest on the right of it. The treasure chest contains a singular quill.
Behind the Giant Tree Between the Two River Paths (1): Behind the big tree that holds up Bat Ship Crazy is a treasure chest that contain one quill. The treasure chest is also in between the two river paths.
Top of Blasto's Hill(1): On the left side (facing away from Hivory Towers) of Shipwreck Creek, an uphill path can be found that leads to Blasto and a treasure chest that has a singular quill inside of it.
Right of the Tree Next to the Rocks with towels (1): The right (facing away from Hivory Towers) of the gray rocks with the Yooka and Laylee towels is giant tree. Past that giant tree is a treasure chest found in the corner. The chest contains a single quill.
Left of the Entrance of Hivory Towers (1): To the left (facing Hivory Towers) of the entrance of Hivory Towers is a small descending path, and a treasure chest can be found on the way down.
Below the Bridge Between Shipwreck Creek and the Entrance of Hivory Towers (1): Below the natural bridge that lies between Shipwreck creek and the Entrance of Hivory Towers is a path that contains a treasure chest containing one quill.
In the Hivory Towers Entrance, found on top of the wall that has fences and acts as a barrier for Shipwreck Creek and the path into Hivory Towers. To reach the top, the player must have the Reptile Roll ability. There's a gold slope located in the "tutorial" section of the path to Hivory Towers. Once they reach the top of the slope, the player can double jump towards the rock cliff, and then using either the air attack or glide to extend the jump. Climb up the rocks and to the right will be the Health Extender.
Inside the nose of the Golden Capital B statue. Use Buddy Slam on top of it to break the nose. Doing so will unlock the Petty Vandalism trophy.